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拟南芥成花调控途径的研究进展.pdf

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拟南芥成花调控途径的研究进展.pdf

<p>J 殮 ·SHANXI AGRIC ， UNIV ·（ &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;） &nbsp; （ &nbsp; &nbsp; &nbsp;） ， （） ： &nbsp; &nbsp; &nbsp; &nbsp;： - &nbsp; &nbsp; ： - &nbsp; &nbsp; ： &nbsp; &nbsp;（-） ， &nbsp;（ &nbsp;） ， &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp;， &nbsp; ， &nbsp; &nbsp; ： &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;* &nbsp; &nbsp; ： &nbsp; &nbsp;， &nbsp; ， &nbsp; &nbsp; &nbsp;，Tel ： ，E-mail ： com &nbsp; &nbsp; ： &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （YBSJJ） ； &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （D-） ； &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （yydzx） &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， ， &nbsp; &nbsp; ， &nbsp; &nbsp; ， &nbsp; &nbsp; ， &nbsp; &nbsp; * （· &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; ； · &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; ） &nbsp; ： &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp;： &nbsp; &nbsp; ； &nbsp; &nbsp; ； &nbsp; &nbsp;； &nbsp; &nbsp; &nbsp; &nbsp;：Q &nbsp; &nbsp; &nbsp; ：A &nbsp; &nbsp; &nbsp;： -（）- doi ： ·j cnki issn- Regulation pathways of flowering in &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;* （ &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp;； &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; - &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp;） Abstract ： ObjectiveTheflowerformationprocessisessentialduringvegetativetoreproductivephasetransition Ap- propriatebloomingtimeiscrucialforcropproductionandbreedingwork MethodsPresentpapersummarizedthere- centinsightsinregulationpathwaysoffloweringinplants &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;ResultsThemechanismofthesix importantfloweringregulatorypathwayswasexplicated ；unsolvedproblemsandpotentialinterestsoffutureresearch werediscussed ConclusionTheinformationpresentedinthepapercansupporttheoreticalbasisofmechanismre- searchoffloweringregulationandartificialfloweringinductiontechnology Keywords ：Controlofflowerformation ，Floweringtime ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;，Gene &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 1 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;： &nbsp; &nbsp; 、 &nbsp; &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp;、 &nbsp; &nbsp; 、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 1·1 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp;FLOWERING LO- CUSC（FLC） &nbsp; 。 &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;： &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp;MADS-box &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; - &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; - &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp;） &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; ） ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp;- &nbsp; &nbsp; &nbsp;（ &nbsp; ） &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp;） &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp;（ &nbsp; &nbsp;） &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; - &nbsp; &nbsp; &nbsp;（ &nbsp; &nbsp;） &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;- &nbsp; - &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp; ）、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;- &nbsp; &nbsp; （ &nbsp; &nbsp;）、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp;） &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp;）、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp;） &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp;LongnoncodingRNAs（lncRNAs） &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; COLDAIR &nbsp;COLDWRAP &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;PROTEASOMEALPHASUB- UNITF（PAF） &nbsp; &nbsp; FRI &nbsp; &nbsp; &nbsp; 。 1·2 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;（ &nbsp; ） &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; PHYTOCHROMEA（PhyA） &nbsp; &nbsp;SPAs- COP &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;CO &nbsp; ， &nbsp; &nbsp; &nbsp; ； &nbsp; ，PhyB &nbsp; &nbsp;CO &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; ，PhyB &nbsp; &nbsp; &nbsp; &nbsp;VOZ &nbsp;VOZ &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ；PhyC &nbsp; &nbsp; &nbsp; &nbsp; PhyA &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ； PhyD 、PhyE &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; CRYPTOCHROME （CRY）、CRY &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp;- &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;（ &nbsp; &nbsp;）、 &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;（ &nbsp; &nbsp;） &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp; &nbsp;） &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; FLOWERINGLOCUST（FT） &nbsp; 。FT &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;（ &nbsp; ） ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 1·3 &nbsp; &nbsp; &nbsp; &nbsp; （gibberellins ，GA） &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp;GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; &nbsp; GA &nbsp;GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp;GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;：GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;（ &nbsp; &nbsp; &nbsp; 、 &nbsp; &nbsp; 、 &nbsp; &nbsp; &nbsp;） &nbsp;GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;（ &nbsp; &nbsp; &nbsp;、 &nbsp; &nbsp;、 &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;） ， 。 &nbsp; &nbsp;、 &nbsp; &nbsp;、 &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; DELLA &nbsp; ， &nbsp; &nbsp; &nbsp;GA &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， 。GA &nbsp; &nbsp;DELLA &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp;S &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; DELLA &nbsp;SPOROCYTELESS（SPL） &nbsp; &nbsp; &nbsp;， &nbsp; MADS-box &nbsp; &nbsp;miR ， &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp;） ，GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; ，GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;、 &nbsp; 、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; 。GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp;，DELLA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; DELLA &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp;，GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;P &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; GA &nbsp; &nbsp;C &nbsp; &nbsp; ， &nbsp; GA &nbsp; &nbsp;， &nbsp; &nbsp; GA-DELLA-AP &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; ， &nbsp; GA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 1·4 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; 、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp;： &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp;）、 &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; - &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp;）、 &nbsp; 、 &nbsp; &nbsp;、 &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp;） &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; RNA &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;。 &nbsp; RNA &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;、 &nbsp; &nbsp;、 &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp;RNA &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp;mRNA &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; ； &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; FLD 、FVE &nbsp;REF ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; ， ；LD &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp;SUPPRESSOR OF FRIGIDA（SUF） &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;- &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;- &nbsp; &nbsp; （ &nbsp; &nbsp; ）、 &nbsp; &nbsp; &nbsp;-、 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;- （） &nbsp; &nbsp; ： &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; （ &nbsp; &nbsp; ） &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; - &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; 。 &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; ， &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp; &nbsp;， &nbsp; &nbsp;</p>

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